when an object is removed (not sent to trash, really deleted), any other object that has the deleted as an embed will generate an exception:

An exception has been thrown during the rendering of a template ("Could not find 'Content' with identifier 'array (
'id' => '1571',
'languages' => NULL,
'versionNo' => NULL,
)'") in "EzPublishCoreBundle::content_fields.html.twig".
500 Internal Server Error - Twig_Error_Runtime
2 linked Exceptions: NotFoundException » NotFoundException »

steps to reproduce:

• update to 5.3.3
• set the environment to "dev"
• edit any object, embed a test article A in the body
• save, exit, remove (not to trash) A
• open the object in frontend in a pure Sf siteaccess
• exception (note, on prod there is just a generic server error 500 message)

note, when removing to trash, issue does not show, probably because of https://jira.ez.no/browse/EZP-23592